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Before the dinosaurs arrived on the scene, the world was a very strange place indeed. Much of the Triassic Period was something of a trial and error period for the forces of evolution, as all manner of new reptile groups took over in the wake of the End Permian Extinction. This, then, was a world filled with giant, alien predators across the seas and land, like nothing else the world had seen at that point - and today, we'll be bringing you face to face with them.

From a marine giant that dispatched its prey with immense jaws, to a crocodile relative bigger than some predatory dinosaurs, these are the terrors of the Triassic.


Written, Researched & Presented by: Thomas McGlynn

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Animals
Transcript
00:00Before the dinosaurs evolved, the world was a very strange place indeed.
00:04Much of the Triassic period was something of a trial and error period for the forces of evolution
00:09and all manner of new reptile groups took over in the wake of the end Permian extinction.
00:14This then was a world filled with giant alien predators across the seas and land
00:19like nothing else the world had seen at that point,
00:21and today we'll be bringing you face to face with them.
00:24From a marine giant that dispatched prey with immense jaws
00:27to a crocodile relative bigger than some predatory dinosaurs,
00:31these are the true terrors of the Triassic period.
00:52Before the dinosaurs evolved to rule their respective environments,
00:56much of the world was ruled by a group called the Rauasuchians.
01:00These often enormous, predatory reptiles are related to modern-day crocodiles,
01:05and while the name of the group is informal and unofficial in the taxonomic sense,
01:09it's still widely used to group together several large Triassic-age Pseudasuchians.
01:13While many Rauasuchians were indeed very large,
01:17none were larger than Fosolosuchus tenax,
01:19a species native to the Los Colorados formation of northwestern Argentina
01:23around 220 million years ago.
01:26Growing to lengths of 8 to 10 meters when fully grown,
01:30Fosolosuchus has earned the title of the largest known
01:32non-dinosaurian terrestrial apex predator.
01:36Fosolosuchus looked much like a laterally flattened crocodile,
01:39with longer legs and a stiff tail,
01:40that allowed it to lead a life away from the water's edge.
01:43The jaws were of course the most impressive feature of this animal,
01:47and were filled with enormous recurved, razor-sharp teeth,
01:49which were used for manipulating and crushing through the bodies of the reptiles and synapsids
01:53that shared its environment.
01:55Unlike others of its kind,
01:57Fosolosuchus possessed one large row of tough, bony osteoderms
02:01that ran along the animal's back,
02:02likely protecting it from the jaws of members of the same species.
02:06Fosolosuchus actually co-existed with some of the first dinosaurs,
02:09and given its size, stature and dental weaponry,
02:12we can assume that it most likely hunted and fed upon them too.
02:15It co-existed with Riocasaurus,
02:18a 10-metre-long prosauropod dinosaur
02:20that could have weighed up to 3,000 kilograms when fully grown.
02:24Fosolosuchus most likely utilised its giant teeth
02:26to slash into the neck and flanks of young or weak Riocasaurus,
02:30sustaining itself on a huge quantity of meat for a considerable period of time.
02:40One of the most well-documented and widespread marine predators of the Triassic period
02:44was the giant ichthyosaur Cimbospondylus.
02:47C. yungorum,
02:48the largest species within the genus
02:50and one of the largest ichthyosaurs known to science at this point in time,
02:54was capable of growing to lengths in excess of 17 metres when fully grown.
02:58That's roughly the same size as a fully grown sperm whale,
03:00a large mosasaurus and the same size as some sauropods.
03:05Cimbospondylus sat within the Cimbospondylidae family of ichthyosaurs
03:09and as a result drastically differed from those you might be familiar with from the Jurassic period.
03:14It evolved the classic large eyes and long snout that were typical for the ichthyosaurs,
03:18but its body was much longer than the likes of ichthyosaurus and ophthalmosaurus.
03:23Moreover, it may have lacked a dorsal fin altogether,
03:25resembling the sea serpent-esque mosasaurs
03:28that would arise in the late Cretaceous period more than anything else.
03:31Giant ichthyosaurs such as Cimbospondylus
03:34were akin to the whales and dolphins of the modern day
03:36and lived rather similar lives,
03:38feeding upon fish, cephalopods, large bivalves and even marine tetrapods.
03:43Cimbospondylus yungorum in particular
03:45is believed to have been an apex predator of medium to large-sized marine reptiles
03:49such as other ichthyosaurs,
03:51potentially including those belonging to the same species or genus.
03:54The bones belonging to Cimbospondylus yungorum were initially discovered
03:58in Nevada's Augusta Mountains by Paul Martin Sander in 1998,
04:02who eventually unearthed them between 2014 and 2015,
04:06naming the species formally in 2021.
04:09The genus as a whole was much more widespread,
04:11spanning Canada, the western United States,
04:14the Caribbean and southern Europe between 251 to 242 million years ago.
04:20Cimbospondylus yungorum is specifically associated with the Favret formation of Nevada,
04:25where it lived alongside up to three other species of the same genus,
04:28C. Petrinus, C. Nicholseye and C. Dwellferi.
04:32The latter three were much smaller than C. Yungorum, of course,
04:35and may have occasionally fallen victim to the jaws of the immense predator
04:38as it passed through shallow coastal waters in search of food.
04:42Cimbospondylus likely hung close to the surface waters across the oceans in which it lived
04:46due to the high quantities of oxygen in contrast with the deeper waters below,
04:50in which very little could survive due to the toxic conditions.
04:59Nothing on Earth today resembles Arizonasaurus.
05:03It looks like it's something between a dinosaur, a crocodilian,
05:07and a Permian-age synapsid such as Dimetrodon,
05:09and yet it's not really closely related to any of these groups.
05:12It is indeed an archosaur, and a pseudosuchion at that,
05:16which places it closer to crocodiles than the synapsids,
05:18but sits within the clad Poposauroidea, an unusual and highly derived group.
05:24Arizonasaurus babetii, as you might have guessed,
05:26resided in Arizona during the Middle Triassic period,
05:29roughly 243 million years ago, and was named by Samuel Paul Wells in 1947.
05:34The skeleton is rather well known,
05:37and one near-complete specimen on Earth in 2002
05:39has allowed scientists to gain a good understanding
05:42of what this animal looked like in life.
05:44Much like Spinosaurus,
05:46Arizonasaurus boasted an impressive sail atop its back
05:49that was made from elongated neural spines.
05:52This structure, much like the ones
05:53harnessed by the likes of Dimetrodon and Adaphosaurus
05:56before it in the Permian,
05:57may have been used to regulate its body temperature
05:59in the hot, arid environments of ancient Nevada's Moenkopee formation,
06:03but it could also have held a function in mating displays or communication.
06:06Large, fully grown Arizonasaurus specimens
06:09could have attained lengths of 3 metres from nose to tail,
06:12and may have weighed between 70 and 100 kilograms at that size.
06:16They were, of course, carnivores,
06:18with recurved serrated teeth lining their powerful jaws
06:20that would have been used to cut into and slash at prey,
06:23slicing through flesh with terrifying efficiency.
06:26When Arizonasaurus was alive,
06:28the Moenkopee formation was characterised
06:30by a vast, arid coastal floodplain,
06:32and an ecosystem that was still in recovery
06:34following the catastrophic events of the end Permian extinction event.
06:38Arizonasaurus most likely hunted in land,
06:40preying upon the tetrapods attempting to scrape a living
06:43in the hot, dry rocks of ancient Arizona.
06:45Its diet may have consisted of
06:47Temnospondyl amphibians,
06:48Rhynchosaurs,
06:49Dicinodonts,
06:50and Trilophosaurids such as Anisodontosaurus.
07:00Erythrosuchus africanus
07:01Erythrosuchus africanus
07:01boasted one of the most impressive skulls
07:03in all of natural history.
07:05As the largest predator of its time,
07:07growing up to lengths of between 4 and 5 metres
07:09when fully grown,
07:10it was an impressive-looking animal,
07:12but almost a quarter of its body length
07:14was made up by an immense
07:15theropod dinosaur-like head.
07:17In life, its post-cranial body
07:19would have resembled something akin
07:20to a modern-day crocodile, with legs positioned
07:23towards the underside of its body
07:24rather than adopting the sprawling,
07:26lizard-like gait seen in most large reptiles of its time.
07:29Within the immense jaws of Erythrosuchus
07:31sat a series of sharp, conical teeth
07:33that would have been used to pierce the flesh of its prey,
07:35causing puncturing and bleeding damage
07:37to the head, neck, and torso.
07:38This particular species was a resident
07:41of the early Middle Triassic period
07:42of what would eventually become South Africa,
07:45living between 249 to 244 million years ago.
07:49It's been known to science since 1905,
07:52and early reconstructions of Erythrosuchus
07:54show an animal that essentially resembled
07:56a quadrupedal Tyrannosaurus rex,
07:58albeit one with a proportionately enormous head.
08:01It too was an archosauroform,
08:03sitting within the family Erythrosuchidae,
08:05where it was the largest of its kind,
08:06and one of the largest terrestrial apex predators
08:08of the entire Triassic period.
08:11But why did Erythrosuchus boast such a large head,
08:14a head that was in fact proportionately larger
08:16than that of any known reptile alive or extinct?
08:19Well, the skull is thought to have been the anchor point
08:21for immense jaw muscles,
08:22which afforded Erythrosuchus
08:24an exceptionally strong bite force.
08:26It likely hunted large, slow-moving animals
08:28that were tough to bring down,
08:30such as the bulky herbivorous Dicinodonts,
08:32and therefore boasting such an impressive
08:34cranial and muscle structure,
08:36with teeth shaped like steak knives,
08:37would have given it the power it needed
08:39to reliably bring down the toughest herbivores of its day.
08:42Despite the impressive size of the skull,
08:44it was relatively lightweight,
08:45with multiple weight-saving air pockets
08:47known as fenestrae lining the bone.
08:49This, when combined with an exceptionally thick
08:51and powerful neck,
08:52meant that Erythrosuchus had no trouble at all
08:54wielding such a skull as its primary weapon.
09:02With size estimates ranging from 7 to 12 meters in length,
09:07Smilosuchus Gregorii was one of the most impressive
09:09predators of the entire Mesozoic.
09:11It too resembled a crocodile,
09:13more so than any of the other entries in today's video,
09:16but it was in no way related.
09:17It was in fact a phytosaur,
09:19a member of a group of basal archosauriforms
09:21that superficially looked like crocodiles.
09:24Its generic name translates into English as knife crocodile,
09:27and given the shape and size of its teeth,
09:29that's not particularly hard to see why.
09:32This species proudly wielded immense tusk-like teeth
09:34at the front of its mouth
09:35for piercing and gripping prey items,
09:38which were then processed closer to the back of the jaws
09:40with blade-like slicing teeth.
09:42Initially described in 1995
09:44from remains that were once thought to belong
09:46to the closely related Leptosuchus,
09:48Smilosuchus lived in what is today Arizona
09:50between 221 and 205 million years ago.
09:54While three species have been named and described,
09:57Smilosuchus Gregorii is the largest,
09:59possessing a large rostral crest atop its snout,
10:02just above those immense tusk-like teeth.
10:04Its remains have been discovered
10:05in Arizona's Chinle Formation,
10:07a particularly fossil-rich site
10:08containing the remains of large reptiles,
10:10therapsids, and of course, dinosaurs.
10:13Smilosuchus was not the only large predator
10:15present in the Chinle Formation at the time it lived,
10:17and it may have faced competition
10:18from its distant terrestrial relative,
10:20Postosuchus.
10:23Smilosuchus is believed to have lain in wait
10:24in rivers and lakes just at the water's edge,
10:27while herbivores such as Placarias,
10:29the giant Dysynodon,
10:30would come down to drink.
10:31At the very last second,
10:33Smilosuchus would emerge,
10:34jaws agape,
10:35dragging its unlucky victim down to a watery grave.
10:38Also living here were innumerable numbers
10:40of Coelophysis,
10:41a primitive dinosaur that is believed
10:43to have lived in large groups,
10:44as well as Dismatosuchus,
10:46a particularly well-armored herbivore
10:48belonging to a group of archosaurs
10:49known as the Aetosaurs.
10:56The early Mesozoic then
10:57was filled with many terrifying
10:59non-dinosaurian threats,
11:01but today we've only really met a few of them.
11:04In the seas,
11:05enormous ichthyosaurs,
11:06much larger than Simba Spondylus,
11:08were beginning to spread out
11:09and diversify towards the end of the Triassic,
11:11while on land,
11:12the very first carnivorous dinosaurian apex predators
11:14such as the mighty Herrerasaurus
11:16were taken over.
11:17By the end of the Triassic,
11:19the stage was set for the dinosaurs
11:20to become the dominant group of reptiles
11:22across the entire world.
11:24Thank you so much as always
11:25for watching to the end of this video,
11:27we really, really appreciate it.
11:29Let us know in the comments
11:30what your favourite Triassic predator was.
11:32Don't forget to like, hype,
11:33and subscribe to Dinosaur Discovery,
11:35and we'll see you next time.

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